With digital-currency trading volume having plunged, some of the biggest crypto exchanges are turning to unorthodox practices to boost activity and win market share.

Bitfinex, FCoin and OKex are encouraging startups to drive depositors to their online-trading platforms to get coins listed. Other exchanges, such as Binance and KuCoin, are embracing listing fees that can differ by project. Many exchanges are also rolling out their own "native coins," which traders then use to vote on potential listings.

The practices are in contrast to those at traditional exchanges, which charge lower flat rates and don’t require issuers to direct trading traffic to the exchange. Other bourses don’t poll existing users on which securities get to trade.

The embrace of the unusual comes as crypto exchanges feel the pain that’s swept through the digital currency market after coin prices collapsed 50 percent on average. Trading volume has plummeted 80 percent since a January peak, according to CoinMarketCap.com data. And new types of coins and exchanges, which are billed as cheaper and easier to use, threaten to eat deeper into fee revenue.

"The market downturn has certainly contributed to an increase in unorthodox strategies by token issuers and exchanges," Lucas Nuzzi, director of technology research at Digital Asset Research, said in an email.

For coin issuers looking to get exposure to investors, the practices can be confounding. Christopher Franco, co-founder of Washington, North Carolina-based blockchain startup Expanse, said KuCoin quoted him a listing fee of 50 Bitcoins -- roughly $315,000 at current prices. Expanse didn’t go through with a listing.

“We can pay for it, but it doesn’t justify the means,” Franco said in a phone interview. Many startups prefer to invest such sums into research and development and marketing.

KuCoin spokesman disputed the 50 Bitcoin listing fee, and said its listing prices vary by startup. The exchange doesn’t disclose them. “The listing fee is not the key factor for listing a project, the project itself is,” spokesman Miles Wu said.

Crytpo exchange fees can be difficult to determine ahead of time, and two startups of the same size applying to list to the same exchange may be charged different rates. In contrast, Nasdaq Inc. charges $50,000 to list a company with up to 15 million shares, and $225,000 to list a company with more than 100 million shares.

The motivation behind the unusual moves is clear enough. Fees have contributed about $1 billion to the exchanges’ revenue to date, according to Lex Sokolin of Autonomous Research. Listing prices at some exchanges in Asia have reached as much as $1 million, according to Michael Jackson, partner at Mangrove Capital Partners.

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